the animal kingdom. Birds, notwithstanding their great locomotive powers, form no exception to the general rules already laid down; but, in this class, as in plants and terrestrial quadrupeds, different groups of species are circumscribed within definite limits. We find, for example, ane assemblage in the Brazils, another in the same latitudes in Central Africa, another in India, and a fourth in New Holland. Of twenty-six different species of land birds found in the Galapagos archipelago, all, with the exception of one, are distinct from those inhabiting other parts of the globe;* and in other archipelagos a single island sometimes contains a species found in no other spot on the whole earth; as is exemplified in some of the parrot tribes. In this extensive family, which are, with few exceptions, inhabitants of tropical regions, the American group has not one in common with the African, nor either of these with the parrots of India.† Another illustration is afforded by that minute and beautiful tribe, the humming-birds. The whole of them are, in the first place, peculiar to the new world; but some species are confined to Mexico, while others exist only in some of the West India Islands, and have not been found elsewhere in the western hemisphere. Yet there are species of this family which have a vast range, as the Trochilus flammifrons (or Mellisuga Kingii), which is found over a space of 2500 miles on the west coast of South America, from the hot dry country of Lima to the humid forests of Tierra del Fuego. Captain King, during his survey in the years 1826-30, found this bird at the Straits of Magellan, in the month of May-the depth of winter-sucking the flowers of a large species of fuchsia, then in bloom, in the midst of a shower of snow. The ornithology of our own country affords one well-known and striking exemplification of the law of a limited specific range; for the common grouse (Tetra scoticus) occurs nowhere in the known world except in the British isles. Some species of the vulture tribe are said to be cosmopolites; and the common wild goose (Anas anser, Linn.), if we may believe some ornithologists, is a general inhabitant of the globe, being met with from Lapland to the Cape of Good Hope, frequent in Arabia, Persia, China, and Japan, and in the American continent from Hudson's Bay to South Carolina. An extraordinary range has also been attributed to the nightingale, which extends from western Europe to Persia, and still farther. In a work entitled Specchio Comparativo,§ by Charles Bonaparte, many species of birds are enumerated as common to Rome and Philadelphia: the greater part of these are migratory, but some of them, such as the long-eared owl (Strix otus), are permanent in both countries. The correspondence of the ornithological fauna of the eastern and western hemispheres increases considerably, as might have been anticipated, in high northern latitudes. * Their facilities of diffusion.-In parallel zones of the northern and southern hemispheres, a great general correspondence of form is observable, both in the aquatic and terrestrial birds; but there is rarely any specific identity; and this phenomenon is truly remarkable, when we recollect the readiness with which some birds, not gifted with great powers of flight, shift their quarters to different regions, and the facility with which others, possessing great strength of wing, perform their aërial voyage. Some migrate periodically from high latitudes, to avoid the cold of winter, and the accompaniments of cold,-scarcity of insects and vegetable food; others, it is said, for some particular kinds of nutriment required for rearing their young: for this purpose they often traverse the ocean for thousands of miles, and recross it at other periods, with equal security. Periodical migrations, no less regular, are mentioned by Humboldt, of many American water-fowl, from one part of the tropics to another, in a zone where there is the same temperature throughout the year. Immense flights of ducks leave the valley of the Orinoco, when the increasing depth of its waters and the flooding of its shores prevent them from catching fish, insects, and aquatic worms. They then betake themselves to the Rio Negro and Amazon, having passed from the eighth and third degrees of north latitude to the first and fourth of south latitude, directing their course south-south-east. In September, when the Orinoco decreases and re-enters its channel, these birds return northwards.t The insectivorous swallows which visit our island would perish during winter, if they did not annually repair to warmer climes. It is supposed that in these aerial excursions the average rapidity of their flight is not less than fifty miles an hour; so that, when aided by the wind, they soon reach warmer latitudes. Spallanzani calculated that the swallow can fly at the rate of ninety-two miles an hour, and conceived that the rapidity of the swift might be three times greater. The rate of flight of the eider duck (Anas mollissima) is said to be ninety miles an hour; and Bachman says that the hawk, wild pigeon (Columba migratoria), and several species of wild ducks, in North America, fly at the rate of forty miles an hour, or nearly a thousand miles in twenty-four hours. § When we reflect how easily different species, in a great lapse of ages, may be each overtaken by gales and hurricanes, and, abandoning themselves to the tempest, be scattered at random through various regions of the earth's surface, where the temperature of the atmosphere, the vegetation, and the animal productions, might be suited to their wants, we shall be prepared to find some species capriciously distributed, and * Bachman, Silliman's Amer. Journ., No. 61, p. 92. Voyage aux Régions Equinoxiales, tome vii. p. 429. Fleming, Phil. Zool., vol. ii. p. 43. Silliman's Amer. Journ., No. 61. p. 83. to be sometimes unable to determine the native countries of each. Captain Smyth informs me, that, when engaged in his survey of the Mediterranean, he encountered a gale in the Gulf of Lyons, at the distance of between twenty and thirty leagues from the coast of France, which bore along many land birds of various species, some of which alighted on the ship, while others were thrown with violence against the sails. In this manner islands become tenanted by species of birds inhabiting the nearest mainland. Geographical Distribution and Dissemination of Reptiles. A few facts respecting the third great class of vertebrated animals will suffice to show that the plan of nature in regard to their location on the globe is perfectly analogous to that already exemplified in other parts of the organic creation, and has probably been determined by similar causes. Habitations of reptiles. Of the great saurians, the gavials which inhabit the Ganges differ from the cayman of America, or the crocodile of the Nile. The monitor of New Holland is specifically distinct from the Indian species; these latter, again, from the African, and all from their congeners in the new world. So in regard to snakes; we find the boa of America represented by the python, a different though nearly allied genus in India. America is the country of the rattlesnake; Africa, of the cerastes; and Asia, of the hooded snake, or cobra di capello. The amphibious genera Siren and Menopoma belong to North America, possessing both lungs and gills, and respiring at pleasure either air or water. The only analogous animal of the old world is the Proteus anguinus of the lakes of Lower Carniola, and the grotto of Adelsberg between Trieste and Vienna.* There is a legend that St. Patrick expelled all reptiles from Ireland; and certain it is that none of the three species of snakes common in England, nor the toad, have been observed there by naturalists. They have our common frog, and our water-newt, and according to Ray (Quad. 264.), the green lizard (Lacerta viridis). Migrations of the larger reptiles.-The range of the large reptiles is, in general, quite as limited as that of some orders of the terrestrial mammalia. The great saurians sometimes cross a considerable tract in order to pass from one river to another; but their motions by land are generally slower than those of quadrupeds. By water, however, they may transport themselves to distant situations more easily. The larger alligator of the Ganges sometimes descends beyond the brackish water of the delta into the sea; and in such cases it might chance to be drifted away by a current, and survive till it reached a shore at some distance; but such casualties are probably very rare. Turtles migrate in large droves from one part of the ocean to another * Richardson, Brit. Assoc. Rep., vol. v. p. 202. during the ovipositing season; and they find their way annually to the island of Ascension, from which the nearest land is about 800 miles distant. Dr. Fleming mentions, that an individual of the hawk's bill turtle (Chelonia imbricata), so common in the American seas, has been taken at Papa Stour, one of the West Zetland Islands; and, according to Sibbald, "the same animal came into Orkney." Another was taken, in 1774, in the Severn, according to Turton. Two instances, also, of the occurrence of the leathern tortoise (C. coriacea), on the coast of Cornwall, in 1756, are mentioned by Borlase. These animals of more southern seas can be considered only as stragglers, attracted to our shores during uncommonly warm seasons by an abundant supply of food, or carried by the Gulf stream, or driven by storms to high latitudes. Some of the smaller reptiles lay their eggs on aquatic plants; and these must often be borne rapidly by rivers, and conveyed to distant regions in a manner similar to the dispersion of seeds before adverted to. But that the larger ophidians may be themselves transported across the seas, is evident from the following most interesting account of the arrival of one at the island of St. Vincent. It is worthy of being recorded, says Mr. Guilding, "that a noble specimen of the Boa constrictor was lately conveyed to us by the currents, twisted round the trunk of a large sound cedar tree, which had probably been washed out of the bank by the floods of some great South American river, while its huge folds hung on the branches, as it waited for its prey. The monster was fortunately destroyed after killing a few sheep, and his skeleton now hangs before. me in my study, putting me in mind how much reason I might have had to fear in my future rambles through the forests of St. Vincent, had this formidable reptile been a pregnant female, and escaped to a safe retreat.” CHAPTER XXXIX. LAWS WHICH REGULATE THE GEOGRAPHICAL DISTRIBUTION OF SPECIES-continued. Geographical distribution and migration of Fish-of Testacea-of Zoophytes— Distribution of Insects-Migratory instincts of some species-Certain types characterize particular countries-Their means of dissemination-Geographical distribution and diffusion of man-Speculations as to the birth-place of the human species-Progress of human population-Drifting of canoes to vast distances-On the involuntary influence of man in extending the range of many other species. Geographical Distribution and Migrations of Fish. ALTHOUGH We are less acquainted with the habitations of marine animals than with the grouping of the terrestrial species before described, yet it Brit. Animals, p. 149., who cites Sibbald. is well ascertained that their distribution is governed by the same general laws. The testimony borne by MM. Péron and Lesueur to this important fact is remarkably strong. These eminent naturalists, after collecting and describing many thousand species of marine animals which they brought to Europe from the southern hemisphere, insist most emphatically on their distinctness from those north of the equator; and this remark they extend to animals of all classes, from those of a more simple to those of a more complex organization—from the sponges and Medusa to the Cetacea. "Among all those which we have been able to examine," say they, "with our own eyes, or with regard to which it has appeared to us possible to pronounce with certainty, there is not a single animal of the southern regions which is not distinguished I y essential characters from the analogous species in the northern seas."* On comparing the freshwater fish of Europe and North America, Sir John Richardson remarks, that the only species which is unequivocally common to the two continents is the pike (Esox lucius); and it is curious that this fish is unknown to the westward of the Rocky Mountains, the very coast which approaches nearest to the old continent. According to the same author the genera of freshwater fish in China agree closely with those of the peninsula of India, but the species are not the same. "As in the distribution," he adds, "of marine fish, the interposition of a continent stretching from the tropics far into the temperate or colder parts of the ocean, separate different ichthyological groups; so with respect to the freshwater species, the intrusion of arms of the sea running far to the northwards, or the interposition of a lofty mountain-chain, effects the same thing. The freshwater fish of the Cape of Good Hope and the South American ones, are different from those of India and China, &c.”‡ Cuvier and Valenciennes, in their "Histoire des Poissons," observe, that very few species of fish cross the Atlantic. Although their statement is correct, it is found that a great many species are common to the opposite sides of the Indian Ocean, inhabiting alike the Red Sea, the eastern coast of Africa, Madagascar, the Mauritius, the Indian Ocean, the southern seas of China, the Malay archipelago, the northern coasts of Australia, and the whole of Polynesia !§ This very wide diffusion, says Sir J. Richardson, may have been promoted by chains of islands running east and west, which are wanting in the deep Atlantic. An archipelago extending far in longitude, favours the migration of fish by multiplying the places of deposit for spawn along the shores of islands, and on intervening coral banks; and in such places, also, fish find their appropriate food. The flying fish are found (some stragglers excepted) only between the tropics in receding from the line, they never approach a higher latitude * Sur les Habitations des Animaux Marins.—Ann. du Mus., tome. xv., cited by Prichard, Phys. Hist. of Mankind, vol. i. p. 51. Brit. Assoc. Reports, vol. v. p. 203. Report to the Brit. Assoc., 1845, p. 192. |